Inocybe ceskae
No common name
Inocybaceae

Species account author: Ian Gibson.
Extracted from Matchmaker: Mushrooms of the Pacific Northwest.

Introduction to the Macrofungi

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Map

E-Flora BC Static Map

Distribution of Inocybe ceskae
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Species Information

Summary:
Inocybe ceskae is characterized by a yellowish beige, straw-coloured to pale ochre, smooth and sticky cap, a thin whitish evanescent veil in young fruitbodies, "shortly heterodiametrical nodulose spores with rather protruding knobs", and short cystidia (averaging <= 50 microns long), (Esteve-Raventos(1)). The whitish stem is pruinose all over and most often has a marginate bulb. This is one of two species of the Inocybe mixtilis group that are known from the Pacific Northwest. It is difficult to distinguish Inocybe ceskae from Inocybe occulta, the one other known PNW species in the group, and the use of the "Inocybe mixtilis group" would generally be appropriate if specimens are not DNA-sequenced. "Typical collections [of I. occulta] are characterized by the heterodiametrical spores, pleurocystidia 34-69 microns long and a pileus of intense colour. Nevertheless, those characters are somewhat overlapping in I. mixtilis, I. ceskae and I. occulta", (Esteve-Raventos(1) with microns designated by mu-m and Latin names italicized). See also SIMILAR section.
Gills:
"nearly free to adnate or adnexed with decurrent tooth", 0.2-0.5cm broad, "ventricose to subventricose", "moderately crowded to rather crowded", 40-60 gills reaching stem, with 1-3 subgills between neighboring gills; initially whitish, beige or grayish, "showing a rather distinctive pinkish or purplish hue", especially toward the gill edge, "then becoming brownish grey or even deep brown to orange or rusty brown when old"; "edge crenulate, whitish when young, sometimes concolorous with age"
Stem:
2.0-4.5cm x 0.2-0.5cm at top, 0.6-0.9cm wide at bulb, cylindric or widening slightly downwards, base most often with an abrupt marginate bulb; stem solid, fibrous; whitish when young, then becoming yellowish when old but not especially browning or darkening; evenly and densely pruinose all over
Veil:
velipellis "distinct in young fruitbodies, as a very thin whitish coating especially at margin" but also over the entire cap, "soon disappearing and nearly absent in washed or aged specimens"; "cortina not seen, even in young primordia"
Odor:
"more or less spermatic when cut, otherwise sometimes indistinctive"
Microscopic spores:
spores (6.7)7-11.1(11.5) x (5)5.2-7.6(7.7) microns, "heterodiametrical to subheterodiametrical, seldom subisodiametrical, often with a curved adaxial face in profile", presenting (8)9-12(14) "distinct and rounded to conical-obtuse knobs, these often irregularly protruding", 0.7-1.4 microns high, 1.1-1.6(2) microns wide at base, "apicula distinct, curved, without germ pore" and sometimes with a large central droplet; basidia (2)4-spored, 18-30 x 7.5-11.5 microns, clavate, sometimes with small droplets, sterigmata 2.5-5(7) microns long; pleurocystidia abundant, metuloid, (38)40-48-60(65) x (16.5)18-22.7-30.5(31) microns, "broadly subutriform with wide neck, but also broadly fusiform, generally not pedicellate or tapering into a short rounded base, rarely with a short pedicel", "rather crystalliferous at apex", walls 2-4.5 microns thick, "but not coalescing at the neck, pale yellowish", cheilocystidia "abundant, similar in shape and size to pleurocystidia", (36)37-46-54(55) x (16)18-22-25(29) microns, "with similar thick walls, rather crystalliferous at apex and sometimes showing a pale yellowish content", gill edge "nearly homogeneous and sterile, composed of numerous cheilocystidia and often rather small subclavate or subovoid paracystidia"; clamp connections present in all tissues
Notes:
Inocybe ceskae occurs in humid and subtemperate coastal regions of the Pacific Northwest (collections from BC and OR) and in one locality in boreal forest in Northern Europe (collections from Finland) (Esteve-Raventos).
EDIBILITY

Habitat and Range

SIMILAR SPECIES
Inocybe occulta is similar: the key says that the cap of I. ceskae is "pale straw-colored, yellowish beige, pale ochre, slightly sticky in wet conditions", and cystidia are <= 50 microns [but range larger], "fusiform, with attenuate but not pedicellate base", whereas the cap of I. occulta is "darker, buff yellow, golden-yellow, orange-yellow or light brown, often with a subhygrophanous appearance", and the cystidia, often >50 microns long, often have "a sublageniform shape, often extending into a distinct neck, though sometimes variable", (Esteve-Raventos(1) who make it clear that characters overlap, as one can see from the detailed descriptions of the cystidia).
Habitat
usually gregarious, rarely single, has been found on the ground in humid and subtemperate coastal regions of the Pacific Northwest, and in boreal forest in Finland

Synonyms

Synonyms and Alternate Names:
Inocybe mixtilis sensu auct. (misapplied name)